X. Read and translate the short text without any dictionary:

Fact of life: Highly sensitive dating techniques tell us that the Earth is between 4.5 and 5.0 thousand million years old. It is generally agreed by scientists that the Earth was originally devoid of life, and that the first living organisms arose by biochemical evolution from complex organic chemicals formed in the atmosphere and seas of early Earth. These first forms of life gave rise to countless millions of species. Most have become extinct, but some have evolved into organisms found today. According to the latest estimates, 20-30 million species share our planet.

XI. Food for thought:

In 1809 Jean-Baptiste de Lamark suggested that the driving force behind evolution was the need for organisms to adapt to changing environmental conditions. His theory became known as the theory of evolution by the inheritance of acquired characteristics. He believed that adaptations developed by an organism during its lifetime could be passed on to its offspring. According to Lamark, modern giraffes might have evolved from a short-necked ancestors in the following way. Giraffes feed on leaves ripped off the branches of trees. When leaves on the lower branches were removed, or when the trees became taller, the ancestral giraffe needed to stretch to reach leaves on higher branches. By continually stretching, their necks lengthened and the ability to grow a slightly longer neck was inherited by the next generation which carried on stretching, and so on.

We know that this explanation of the evolution of the giraffe’s neck is untrue because activities such as stretching to feed do not affect the gametes. Therefore, this type of characteristic acquired during the life of an organism is not inherited by its offspring. Expressed in modern terms, Lamarckism would mean that changes in phenotype could determine the genotype of future generations. This does not agree with modern genetics, and there are no generally accepted examples of acquired characteristics being inherited. Suggest a neo-Darwinian explanation for the evolution of the modern long-necked giraffe from a short-necked ancestor.

Text 6.2. Natural Selection

■ Essential targets:

By the end of this text you should be able to:

· explain what is meant by “ survival of the fittest”;

· distinguish between directional selection, stabilising selection, disruptive selection.

Pre-reading

■ With a partner consider the following questions and try to answer them. Then scan the text to check your answers.

1. What is natural selection?

2. What environmental factors effect on surviving and producing offspring?

■ Read the given text and make your essential assignments:

Survival of the fittest

Darwin had the idea that natural selection is the mechanism that drives evolution after reading An Essay on the Principal of Population by Thomas Malthus, a clergyman and political economist. Malthus argued that, in time, the growth of human populations will outstrip the food supply, and that this will lead to “famine, pestilence, and war”. Darwin applied this idea to populations of other animals and of plants. In his book on the origin of species, Darwin wrote: “There is no exception to the rule that every organic being naturally increases at so high a rate that if not destroyed, the Earth would soon be covered by the progeny of a single pair”. In spite of reproducing quickly, no single species has completely over-run the planet, although the populations of some species may be increasing at any one particular time. Darwin concluded that populations are kept in check by a “struggle for existence” as they compete for limited resources and are exposed to disease. Environmental factors that keep populations in check are called selection pressuresor environmental resistances. These include:

· disease

· competition for resources such as food and a place in which to live

· predation

· lack of light, water, or oxygen

· changes in temperature.

Those organisms best suited to the environmental conditions, with

characteristics that give them an advantage in the “struggle for existence”, will have the best chance of surviving and producing offspring. Their high natality (birth rate) gives them a selective advantage. On the other hand, those will unfavourable characteristics are more likely to die. Their high mortality (death rate) gives them a selective disadvantage. Darwin argued that this difference in natality and mortality results in natural selection. As environmental conditions change certain characteristics within a randomly varying population are favoured, and natural selection occurs. This has become known as the “survival of the fittest”.

In evolution, fitnessis defined as the ability of an organism to pass on its alleles to subsequent generations, compared with other individuals of the same species. The “fittest” individual in a population is the one that produces the largest number of offspring that survive to reproduce themselves. Natural selection by “survival of the fittest” means that the genetic characteristics of a population gradually change from generation to generation in response to changes in the environment. As we shall see in the following spreads, natural selection affects a gene pool by increasing the frequency of alleles that give an advantage, and reducing the frequency of alleles that give a disadvantage. (A gene pool is all the genes and their different alleles present in an interbreeding population.)

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